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How come the Land Cruiser requires high octane gas, yet the 4runner (also hybrid and with same engine) doesn't?

I read in another thread that A-Trac is now automatic in the 6G. If this is true, what turns it on? 4Lo? Crawl mode? MTS? Specifically on the TRD Pro as that's what I'm planning on purchasing. Been trying to find a more detailed explanation and can't find a detailed answer anywhere...

Looking to purchase a 2025 4Runner TRD Pro (put a deposit down) and wanted to see people's opinions on the stock Toyo Tires. How do they compare to the highly rated BF KO3s & the Falcon Wildpeak AT4s? I'm mainly curious in how they compare regarding pavement road noise, but other things like...